Episode 7.3 · 15 min

the first stone tools, the first members of our genus Homo, and the big-toothed Paranthropus

Who made the first stone tools—and what did stone change for our early relatives? From Kenya’s 3.3-million-year-old cores and flakes to hippo butchery at Nyayanga, this episode follows the uncertain beginnings of deliberate tool use, the emergence of Homo, and Paranthropus’s very different answer to changing foods: enormous chewing teeth. By two million years ago, several branches still overlapped, as a taller Homo approached.

Watch this film on YouTube ↗

Visual credits

34 sources

Sources and credits for imagery used in this film, including material incorporated into explanatory animations.

  • Paranthropus aethiopicus: the Black Skull

    Akrasia25

    Resized and reframed; animated crest highlight added by this channel.

  • Stone tool use by a capuchin monkey

    Tiago Falótico

    Resized for video.

  • Paranthropus boisei skull cast, Springfield Science Museum

    Daderot

    Resized for video.

  • Paranthropus boisei skull cast, MGL 95211

    Rama

    Resized and composited; animated cheekbone highlights added by this channel.

  • Young chimpanzee Shangwe cracking a nut at Artis Royal Zoo

    Kitty Terwolbeck

    Resized for video.

  • Paranthropus boisei lower jaw cast, ER 729

    Bjoertvedt

    Resized for video.

  • OH 80 Paranthropus boisei teeth, Figure 1

    Photographs of individual teeth by J Trueba and MJ Ortega; composition by JL Heaton

    Resized for video.

    From Domínguez-Rodrigo et al., First Partial Skeleton of a 1.34-Million-Year-Old Paranthropus boisei from Bed II, Olduvai Gorge, Tanzania, PLOS ONE (2013), doi:10.1371/journal.pone.0080347. Copyright © 2013 Domínguez-Rodrigo et al.

  • Flint-knapping demonstration

    Tonto National Monument

    Resized for video.

    NPS photo.

  • Homo habilis skull, KNM-ER 1813

    Akrasia25

    Resized for video.

  • Paranthropus boisei tooth microwear, Figure 1

    Peter S. Ungar, Frederick E. Grine and Mark F. Teaford

    Figure extracted and resized for video.

    Copyright © 2008 Ungar et al.

  • Flaked and pounded stone tools from Nyayanga, Figure 2

    Isabella Caricola, Luigi Germinario, Emma M. Finestone, Claudio Mazzoli, Laura Bishop, James S. Oliver, Rahab N. Kinyanjui, Peter W. Ditchfield, Richard Potts, Cristina Lemorini and Thomas W. Plummer

    Figure extracted and resized for video.

    © The Author(s) 2026.

  • Lucy skeleton cast

    120

    Resized for video; source image already has a digitally prepared black background.

  • Hadar landscape, Ethiopia, 1973

    Raymonde Bonnefille

    Resized for video.

    Fonds Raymonde Bonnefille, MédiHAL hal-05354623; IRD / CNRS / INSU.

  • LD-350-1 mandible drawing

    Brobt

    Rasterized, resized and composited; animated jaw and tooth-row highlights added by this channel.

  • Flakes produced during capuchin nut cracking, Figure 4A–F

    Tomos Proffitt, Paula de Sousa Medeiros, Waldney Pereira Martins and Lydia V. Luncz

    Cropped to Figure 4 panels A–F and resized for video.

  • Oldowan stone tools at the Timiryazev Museum

    Александр Сигачёв

    Cropped to the seven displayed stones, padded and resized for video.

  • Cut marks on a Nyayanga hippo tibia, Figure 3A

    Thomas W. Plummer, James S. Oliver, Emma M. Finestone, Peter W. Ditchfield, Laura C. Bishop, Scott A. Blumenthal, Cristina Lemorini, Isabella Caricola, Shara E. Bailey, Andy I. R. Herries, Jennifer A. Parkinson, Elizabeth Whitfield, Fritz Hertel, Rahab N. Kinyanjui, Thomas H. Vincent, Youjuan Li, Julien Louys, Stephen R. Frost, David R. Braun, Jonathan S. Reeves, Emily D. G. Early, Blasto Onyango, Raquel Lamela-Lopez, Frances L. Forrest, Huaiyu He, Timothy P. Lane, Marine Frouin, Sébastien Nomade, Evan P. Wilson, Simion K. Bartilol, Nelson Kiprono Rotich and Richard Potts

    Cropped to Figure 3A and resized for video.

    Copyright © 2023 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works.

  • Cut mark on a Nyayanga hippo rib, Figure 3B

    Thomas W. Plummer, James S. Oliver, Emma M. Finestone, Peter W. Ditchfield, Laura C. Bishop, Scott A. Blumenthal, Cristina Lemorini, Isabella Caricola, Shara E. Bailey, Andy I. R. Herries, Jennifer A. Parkinson, Elizabeth Whitfield, Fritz Hertel, Rahab N. Kinyanjui, Thomas H. Vincent, Youjuan Li, Julien Louys, Stephen R. Frost, David R. Braun, Jonathan S. Reeves, Emily D. G. Early, Blasto Onyango, Raquel Lamela-Lopez, Frances L. Forrest, Huaiyu He, Timothy P. Lane, Marine Frouin, Sébastien Nomade, Evan P. Wilson, Simion K. Bartilol, Nelson Kiprono Rotich and Richard Potts

    Cropped to Figure 3B and resized for video.

    Copyright © 2023 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works.

  • Paranthropus molars from Nyayanga, Figure 4A–B

    Thomas W. Plummer, James S. Oliver, Emma M. Finestone, Peter W. Ditchfield, Laura C. Bishop, Scott A. Blumenthal, Cristina Lemorini, Isabella Caricola, Shara E. Bailey, Andy I. R. Herries, Jennifer A. Parkinson, Elizabeth Whitfield, Fritz Hertel, Rahab N. Kinyanjui, Thomas H. Vincent, Youjuan Li, Julien Louys, Stephen R. Frost, David R. Braun, Jonathan S. Reeves, Emily D. G. Early, Blasto Onyango, Raquel Lamela-Lopez, Frances L. Forrest, Huaiyu He, Timothy P. Lane, Marine Frouin, Sébastien Nomade, Evan P. Wilson, Simion K. Bartilol, Nelson Kiprono Rotich and Richard Potts

    Cropped to Figure 4A–B, padded and resized for video.

    Copyright © 2023 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works.

  • Paranthropus boisei hand, KNM-ER 101000, Figure 3

    Carrie S. Mongle, Caley M. Orr, Matthew W. Tocheri, Thomas Cody Prang, Frederick E. Grine, Craig Feibel, Biren A. Patel, Olivia Laureijs, Tara E. Hobbs, Stephanie Maiolino, James Rossie, Winfred Mbogo, Anton Du Plessis, John Lonyericho, William Woto Huka, Hilary Sale, Abdi Umuro, Peter Yirgudo, Ibrae Dalacha, Martin Kirinya, Ezekiel Linga, Richard Loki, Apolo Longaye, Malmalo Longaye, Emmanuel Lonyericho, Iyole Loyapan, Nyiber Nakudo, Cyprian Nyete, Meave G. Leakey and Louise N. Leakey; hand outline by Francis Tre Lawrence

    Cropped to the palmar view in Figure 3, padded and resized for video.

    © The Author(s) 2025.

  • Quartz stone tools from Nyayanga, Figure 3

    Emma M. Finestone, Thomas W. Plummer, Peter W. Ditchfield, Jonathan S. Reeves, David R. Braun, Simion K. Bartilol, Nelson Kiprono Rotich, Laura C. Bishop, James S. Oliver, Rahab N. Kinyanjui, Michael D. Petraglia, Paul S. Breeze, Cristina Lemorini, Isabella Caricola, Paul Owich Obondo and Richard Potts

    Cropped to the vein-quartz artifact panel in Figure 3 and resized for video.

    Copyright © 2025 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works. Distributed under a Creative Commons Attribution License 4.0 (CC BY).

  • Nyayanga stone tools by geological source, Figure 3

    Emma M. Finestone, Thomas W. Plummer, Peter W. Ditchfield, Jonathan S. Reeves, David R. Braun, Simion K. Bartilol, Nelson Kiprono Rotich, Laura C. Bishop, James S. Oliver, Rahab N. Kinyanjui, Michael D. Petraglia, Paul S. Breeze, Cristina Lemorini, Isabella Caricola, Paul Owich Obondo and Richard Potts

    Cropped to the Nyanzian, Bukoban and Homa Mountain columns of Figure 3 and resized for video.

    Copyright © 2025 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works. Distributed under a Creative Commons Attribution License 4.0 (CC BY).

  • Long-tailed macaque using a stone to crack a snail

    Lydia V Luncz, Amanda Tan, Michael Haslam, Lars Kulik, Tomos Proffitt, Suchinda Malaivijitnond and Michael Gumert

    Cropped to panel A and resized for video.

    From Resource depletion through primate stone technology, eLife 2017;6:e23647, doi:10.7554/eLife.23647.013.

  • Macaque pounding stone and broken shells

    Lydia V Luncz, Amanda Tan, Michael Haslam, Lars Kulik, Tomos Proffitt, Suchinda Malaivijitnond and Michael Gumert

    Cropped to panel C and resized for video.

    From Resource depletion through primate stone technology, eLife 2017;6:e23647, doi:10.7554/eLife.23647.013.

  • Bokol Dora stone flake, Ethiopia

    David R. Braun

    Cropped from the source image and resized for video.

  • Oldowan chopper from Melka Kunture, Ethiopia

    Didier Descouens

    Cropped to a single artifact view and resized for video.

  • Lomekwi 3 stone core, Figure 4b

    Sonia Harmand, Jason E. Lewis, Craig S. Feibel, Christopher J. Lepre, Sandrine Prat, Arnaud Lenoble, Xavier Boës, Rhonda L. Quinn, Michel Brenet, Adrian Arroyo, Nicholas Taylor, Sophie Clément, Guillaume Daver, Jean-Philip Brugal, Louise Leakey, Richard A. Mortlock, James D. Wright, Sammy Lokorodi, Christopher Kirwa, Dennis V. Kent and Hélène Roche

    Cropped to Figure 4b and resized for video.

    © 2015 Macmillan Publishers Limited. All rights reserved.

  • Captive chimpanzees cracking nuts, S2 Video

    Arroyo A, Hirata S, Matsuzawa T and de la Torre I

    Transcoded, excerpted and resized for video.

    S2 Video from Nut Cracking Tools Used by Captive Chimpanzees (Pan troglodytes) and Their Comparison with Early Stone Age Percussive Artefacts from Olduvai Gorge, PLOS ONE (2016), doi:10.1371/journal.pone.0166788.

  • Gelada at Lalibela, Ethiopia

    Jenn

    Transcoded, excerpted and resized for video.

  • Nyayanga stone-source map, adapted from Figure 2

    Emma M. Finestone, Thomas W. Plummer, Peter W. Ditchfield, Jonathan S. Reeves, David R. Braun, Simion K. Bartilol, Nelson Kiprono Rotich, Laura C. Bishop, James S. Oliver, Rahab N. Kinyanjui, Michael D. Petraglia, Paul S. Breeze, Cristina Lemorini, Isabella Caricola, Paul Owich Obondo and Richard Potts; redraw and animation by AgentHouse

    Geography measured from Figure 2, simplified and redrawn; distance circle and animated highlights added by AgentHouse.

    Copyright © 2025 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works. Distributed under a Creative Commons Attribution License 4.0 (CC BY).

  • Earth normal map

    Solar System Scope

    Converted to relief shading, reprojected and composited into the Africa maps.

    Textures by Solar System Scope (solarsystemscope.com), CC BY 4.0

  • Natural Earth country boundaries, 1:50 million

    Natural Earth; Tom Patterson, Nathaniel Vaughn Kelso and contributors

    Reprojected and styled as coastlines, land fill and country outlines in the Africa map.

  • Natural Earth country boundaries, 1:50 million

    Natural Earth; Tom Patterson, Nathaniel Vaughn Kelso and contributors

    Reprojected and styled as coastlines, land fill and country outlines in the Africa map.

  • Natural Earth country boundaries, 1:50 million

    Natural Earth; Tom Patterson, Nathaniel Vaughn Kelso and contributors

    Reprojected and styled as coastlines, land fill and country outlines in the Africa map.

Stock footage

18 items

Licensed stock footage used in this film, with each item’s creator and source page.

Research references

Selected sources from the episode’s research and scientific figures.

  1. Nature, doi:10.1038/nature14464 ↗
  2. PaleoAnthropology 46–53 ↗
  3. doi:10.1016/j.quaint.2025.110078 ↗
  4. doi:10.1038/nature20112 ↗
  5. full text ↗
  6. doi:10.1073/pnas.2420067122 ↗
  7. Smithsonian early toolkit ↗
  8. Bril et al., functional mastery of percussion ↗
  9. article Fig.2 ↗
  10. 2023 Science ↗
  11. 2025 transport article ↗
  12. doi:10.1038/nature16990 ↗
  13. doi:10.1126/science.aaa1343 ↗
  14. comment doi:10.1126/science.aab0591 ↗
  15. response doi:10.1126/science.aab1122 ↗
  16. primary ↗
  17. OH 80 study, doi:10.1371/journal.pone.0080347 ↗
  18. doi:10.1073/pnas.1104627108 ↗
  19. 2008 PLoS ONE ↗
  20. doi:10.1038/s41586-025-09826-x ↗
  21. doi:10.1038/s41586-025-09594-8 ↗
  22. doi:10.1073/pnas.2530996123 ↗
  23. primary chronology ↗
  24. 2025 Namorotukunan Fig.1 ↗
  25. 9XS8YBP ↗
  26. 2RTM3WL ↗
  27. 8NMMJKX ↗
  28. WENGTYL ↗
  29. y2ihHOCKxUc ↗
  30. doi:10.1163/21915784-20190006 ↗
  31. What is ‘in situ’? ↗
  32. doi:10.1073/pnas.0607909104 ↗
  33. Braun et al. 2016 Oldowan reduction study ↗
  34. exhibit script ↗
  35. doi:10.1126/science.1261735 ↗
  36. Early knapping techniques do not necessitate cultural transmission ↗
  37. PMC ↗
  38. author manuscript ↗
  39. doi:10.1073/pnas.1820177116 ↗
  40. doi:10.1371/journal.pone.0062174 ↗
  41. doi:10.1073/pnas.1222559110 ↗
  42. doi:10.1126/science.adq7315 ↗
  43. published abstract ↗
  44. 2025 commentary, doi:10.17159/sajs.2025/21847 ↗
  45. doi:10.1038/s41467-023-44375-9 ↗
  46. doi:10.1038/nature14224 ↗
  47. doi:10.1002/ar.70100 ↗
  48. published record ↗
  49. PDF ↗
  50. Paranthropus robustus tooth chipping patterns do not support regular hard food mastication ↗
  51. Dietary Adaptation in Paranthropus robustus Postcanine Teeth ↗
  52. Smithsonian ↗
  53. original diagnosis ↗
  54. Granger 2022 ↗
  55. Frost 2022 ↗
  56. museum source ↗
  57. Smithsonian ↗
  58. Smithsonian ↗
  59. institutional main PDF ↗
  60. Oxford repository copy ↗
  61. Figshare metatarsal-analysis script ↗
  62. Lewis/Harmand response ↗
  63. Springfield OH 5 cast ↗
  64. KNM-ER 1813 skull ↗
  65. Lucy blackbg ↗
  66. Bokol Dora Stone Tool ↗
  67. Chimpanzees demonstrating how they use tools ↗
  68. Nature Picture Library originator lead on Vimeo ↗
  69. WRH3ND2 ↗
  70. EemvSe3uMlc ↗
  71. YXMZBMR ↗
  72. Envato excavation category ↗
  73. transport-study release ↗
  74. https://commons.wikimedia.org/wiki/File:Australophithecus_boisei_(cast),_Olduvai_Gorge_-_Springfield_Science_Museum_-_Springfield,_MA_-_DSC03368.JPG ↗

Music

  • Original score
    AgentHouse · Project-made score (From Cosmos to Civilization score catalog v4 with bespoke opening and closing)
    Impulse responses: Detmold University of Music SRIR database, CC BY 4.0.
Footage and image usage

Credits and usage times retained from the published edition or its production credit record. Where present, times identify visual appearances, not documentary chapters. Source and licence corrections are listed in the visual credits above.

  • 120
  • Akrasia25 - source: Original source ↗
  • Arroyo A et al. - source: Original source ↗
  • Bjoertvedt
  • Brobt - source: Original source ↗
  • Carrie S. Mongle et al.; hand outline by Francis Tre Lawrence
  • Daderot
  • David R. Braun
  • Didier Descouens
  • Emma M. Finestone et al.
  • Emma M. Finestone et al.; redraw and animation by AgentHouse - source: Original source ↗
  • Isabella Caricola et al.
  • Jenn
  • Kitty Terwolbeck
  • Lydia V Luncz et al.
  • Natural Earth; Tom Patterson, Nathaniel Vaughn Kelso and contributors - source: Original source ↗
  • Peter S. Ungar, Frederick E. Grine and Mark F. Teaford
  • Photographs of individual teeth by J Trueba and MJ Ortega; composition by JL Heaton
  • Rama - source: Original source ↗
  • Raymonde Bonnefille
  • Solar System Scope - source: Original source ↗
  • Sonia Harmand et al.
  • Thomas W. Plummer et al.
  • Tiago Falótico
  • Tomos Proffitt et al.
  • Tonto National Monument
  • Александр Сигачёв
  • Original animations and figures created by this channel.
  • 120 — 2:01-2:10
  • Akrasia25 — 9:59-10:02, 11:28-11:40
  • allenum — 12:38-12:45
  • AmazingAerial — 3:23-3:29, 6:09-6:13, 12:19-12:26
  • Arroyo A, Hirata S, Matsuzawa T and de la Torre I — 3:45-3:49, 3:49-3:53, 3:53-3:58
  • artush — 12:45-12:50
  • ATWStock — 8:25-8:29
  • Bjoertvedt — 1:36-1:42
  • BlackBoxGuild — 0:51-0:59, 1:21-1:27, 2:14-2:18, 3:58-4:01, 4:56-5:03, 5:10-5:15, 6:23-6:26, 6:26-6:30, 7:04-7:07, 12:12-12:19
  • Brobt — 9:24-9:35, 9:35-9:47, 10:07-10:11
  • Carrie S. Mongle, Caley M. Orr, Matthew W. Tocheri, Thomas Cody Prang, Frederick E. Grine, Craig Feibel, Biren A. Patel, Olivia Laureijs, Tara E. Hobbs, Stephanie Maiolino, James Rossie, Winfred Mbogo, Anton Du Plessis, John Lonyericho, William Woto Huka, Hilary Sale, Abdi Umuro, Peter Yirgudo, Ibrae Dalacha, Martin Kirinya, Ezekiel Linga, Richard Loki, Apolo Longaye, Malmalo Longaye, Emmanuel Lonyericho, Iyole Loyapan, Nyiber Nakudo, Cyprian Nyete, Meave G. Leakey and Louise N. Leakey; hand outline by Francis Tre Lawrence — 13:01-13:12
  • Daderot — 11:58-12:12
  • David R. Braun — 4:24-4:31
  • Didier Descouens — 14:16-14:25
  • DovidPro — 8:18-8:25
  • Emma M. Finestone, Thomas W. Plummer, Peter W. Ditchfield, Jonathan S. Reeves, David R. Braun, Simion K. Bartilol, Nelson Kiprono Rotich, Laura C. Bishop, James S. Oliver, Rahab N. Kinyanjui, Michael D. Petraglia, Paul S. Breeze, Cristina Lemorini, Isabella Caricola, Paul Owich Obondo and Richard Potts — 0:59-1:02, 6:30-6:34, 7:07-7:15
  • Emma M. Finestone, Thomas W. Plummer, Peter W. Ditchfield, Jonathan S. Reeves, David R. Braun, Simion K. Bartilol, Nelson Kiprono Rotich, Laura C. Bishop, James S. Oliver, Rahab N. Kinyanjui, Michael D. Petraglia, Paul S. Breeze, Cristina Lemorini, Isabella Caricola, Paul Owich Obondo and Richard Potts; redraw and animation by AgentHouse — 7:15-7:20
  • Isabella Caricola, Luigi Germinario, Emma M. Finestone, Claudio Mazzoli, Laura Bishop, James S. Oliver, Rahab N. Kinyanjui, Peter W. Ditchfield, Richard Potts, Cristina Lemorini and Thomas W. Plummer — 6:51-7:04
  • Jenn — 12:50-13:01
  • Kitty Terwolbeck — 4:01-4:05
  • Lydia V Luncz, Amanda Tan, Michael Haslam, Lars Kulik, Tomos Proffitt, Suchinda Malaivijitnond and Michael Gumert — 4:09-4:16, 4:22-4:24
  • Natural Earth; Tom Patterson, Nathaniel Vaughn Kelso and contributors — 2:10-2:14, 6:05-6:09, 9:11-9:19
  • panophotograph — 8:51-8:59
  • Peter S. Ungar, Frederick E. Grine and Mark F. Teaford — 12:26-12:33
  • Photographs of individual teeth by J Trueba and MJ Ortega; composition by JL Heaton — 11:21-11:28
  • Rama — 11:40-11:44
  • Raymonde Bonnefille — 9:19-9:24
  • Renaud83 — 4:50-4:56
  • Solar System Scope — 2:10-2:14, 6:05-6:09, 9:11-9:19
  • Sonia Harmand, Jason E. Lewis, Craig S. Feibel, Christopher J. Lepre, Sandrine Prat, Arnaud Lenoble, Xavier Boës, Rhonda L. Quinn, Michel Brenet, Adrian Arroyo, Nicholas Taylor, Sophie Clément, Guillaume Daver, Jean-Philip Brugal, Louise Leakey, Richard A. Mortlock, James D. Wright, Sammy Lokorodi, Christopher Kirwa, Dennis V. Kent and Hélène Roche — 3:29-3:35
  • Thomas W. Plummer, James S. Oliver, Emma M. Finestone, Peter W. Ditchfield, Laura C. Bishop, Scott A. Blumenthal, Cristina Lemorini, Isabella Caricola, Shara E. Bailey, Andy I. R. Herries, Jennifer A. Parkinson, Elizabeth Whitfield, Fritz Hertel, Rahab N. Kinyanjui, Thomas H. Vincent, Youjuan Li, Julien Louys, Stephen R. Frost, David R. Braun, Jonathan S. Reeves, Emily D. G. Early, Blasto Onyango, Raquel Lamela-Lopez, Frances L. Forrest, Huaiyu He, Timothy P. Lane, Marine Frouin, Sébastien Nomade, Evan P. Wilson, Simion K. Bartilol, Nelson Kiprono Rotich and Richard Potts — 6:34-6:37, 6:37-6:46, 7:49-7:57
  • Tiago Falótico — 4:05-4:09
  • Tomos Proffitt, Paula de Sousa Medeiros, Waldney Pereira Martins and Lydia V. Luncz — 4:16-4:22
  • Tonto National Monument — 5:30-5:32
  • Александр Сигачёв — 5:44-5:58

How this film was made

Created and editorially directed by Anders Lunde using a custom production system developed over six months. AI assists research, drafting, programming, and visual production; I review and edit the material and make the final editorial decisions. Narration uses text-to-speech. Visuals combine custom-built scientific animations, AI-generated reconstructions, and credited scientific imagery.

Generated reconstructions illustrate scientific interpretations. They are distinct from photographs, specimens, and measured data.

More about the creator and production →