Episode 6.2 · 17 min

Our Ancestors and the Great Dying

How did Permian therapsids begin acquiring features that mammals would later inherit—and how did any survive the Great Dying? This episode follows their changing teeth, jaws, mouths and limbs through the Karoo, then traces how the Siberian Traps helped trigger heat, oxygen loss and acidification that erased most marine species and devastated life on land.

Watch this film on YouTube ↗

Visual credits

22 sources

Sources and credits for imagery used in this film, including material incorporated into explanatory animations.

  • Madumabisa NHMUK PV R 37363, ventral view

    The Trustees of the Natural History Museum, London

    Cropped/reframed and incorporated into an original animation.

  • Odontocyclops fossil skull

    Nkansahrexford

    Cropped to the skull and incorporated into an original animation.

  • Diictodon spiral burrow cast

    Nkansahrexford

    Cropped/reframed and incorporated into an original animation.

  • Two Diictodon skeletons in a burrow cast

    Nkansahrexford

    Cropped to the two skeletons and incorporated into an original animation.

  • Gorynychus masyutinae skull and interpretive drawing

    Christian F. Kammerer

    Cropped/reframed and annotated with color overlays in the animation.

  • Procynosuchus delaharpeae jaw

    Ghedoghedo

    Cropped/reframed and incorporated into an original animation.

  • Lystrosaurus georgi skeleton

    Ghedoghedo

    Cropped/reframed and incorporated into an original animation.

  • Hill at Karoo National Park

    flowcomm

    Cropped/reframed and incorporated into an original animation.

  • Putorana Plateau, Siberia

    European Space Agency

    Cropped/reframed, including a tighter valley crop, and animated with a slow push-in.

  • Glossopteris fossil leaves in claystone

    James St. John

    Cover-cropped and incorporated into an original animation.

  • Early snow on the Putorana Plateau, Aqua/MODIS, 25 September 2003

    Jeff Schmaltz, MODIS Land Rapid Response Team, NASA GSFC

    Cropped/reframed and animated with a slow push-in.

    NASA should be acknowledged as the source of the material.

  • Thrinaxodon fossil

    Nkansahrexford

    Cropped/reframed and incorporated into an original animation.

  • Proterosuchus fergusi skull (SAM-PK-11208)

    Nkansahrexford

    Cropped/reframed and animated with a slow push-in.

  • Nile crocodile skull

    Zachi Evenor

    Cropped to the crocodile skull and incorporated into an original animation.

  • Dimetrodon grandis skull

    Daderot

    Cropped and traced by AgentHouse for the comparison.

  • Inostrancevia skull

    Ghedoghedo

    Cropped at the bottom and annotated within the animation.

  • Monitor Lizard Roaming in Sungei Buloh

    Anish Sheela

    Transcoded, excerpted, trimmed, reframed, and incorporated into an original animation.

  • A Cold Beverage in a Glass

    Engin Akyurt

    Excerpted, trimmed, reframed, and incorporated into an original animation.

  • PALEOMAP PaleoDEM: 250 Ma reconstruction

    Scotese and Wright (2018)

    Processed into derived map graphics for the nominal 250 Ma reconstruction.

  • PALEOMAP PaleoDEM: 255 Ma reconstruction

    Scotese and Wright (2018)

    Processed into derived globe and map graphics, including coastlines, relief, and a derived river network, for the nominal 255 Ma reconstruction.

  • Eoraptor lunensis skeleton

    Esv - Eduard Solà Vázquez

    Used as an anatomical reference through generated Eoraptor reconstruction imagery and animation.

  • Lufengosaurus huenei holotype mount (IVPP)

    Jonathan Chen

    Used as an anatomical reference through generated Lufengosaurus-like reconstruction imagery and animation.

Research references

Selected sources from the episode’s research and scientific figures.

  1. PLOS Biology paper ↗
  2. Gigliotti et al. 2026 ↗
  3. Huttenlocker et al. 2021 ↗
  4. Norton et al. 2023, craniodental correlates ↗
  5. Rawson et al. 2024 ↗
  6. 2021 fossil-record review ↗
  7. Bishop & Pierce 2024 ↗
  8. Rey et al. 2017, oxygen isotopes ↗
  9. Araújo et al. 2022, inner-ear biomechanics ↗
  10. Martinez et al. 2023, maxilloturbinals ↗
  11. Day et al. 2015 ↗
  12. 2023 Current Biology paper ↗
  13. Brant & Sidor 2024 ↗
  14. DOI: 10.1186/s13358-025-00348-7 ↗
  15. 2021 burrow study ↗
  16. Marine chronology ↗
  17. Primary paper ↗
  18. 2020 paper ↗
  19. Svensen and colleagues 2018 ↗
  20. Cui et al. 2021 ↗
  21. Penn, Deutsch, Payne & Sperling 2018 ↗
  22. Jurikova et al. 2020 ↗
  23. PubMed 42406958 ↗
  24. Joachimski et al. 2012 ↗
  25. 2023 mercury-isotope study ↗
  26. 2023 pollen-chemistry study ↗
  27. formal technical comment ↗
  28. response ↗
  29. Stanley 2016 ↗
  30. DOI: 10.1038/s41467-020-15243-7 ↗
  31. DOI: 10.1130/B35830.1 ↗
  32. DOI: 10.1130/B38563.1 ↗
  33. Primary paper ↗
  34. Fielding et al. 2019 ↗
  35. Wu et al. 2024 comparison ↗
  36. Chu 2025 ↗
  37. Primary paper ↗
  38. Open paper and downloadable supplementary datasets ↗
  39. Huttenlocker 2014 ↗
  40. Primary paper, *Did gorgonopsians survive the end-Permian extinction?* ↗
  41. EarthByte resource ↗
  42. Wang & Wang 2007 ↗
  43. Primary crinoid study and phylogenetic context ↗
  44. Primary paper ↗
  45. Chen & Benton 2012 ↗
  46. PLOS article ↗
  47. author-linked code deposit ↗
  48. ESA / Copernicus, Putorana Plateau, 2016 ↗
  49. NASA Earth Observatory, Early Snow on Russia's Putorana Plateau ↗
  50. IUGS Meishan geoheritage page ↗
  51. TimeScale Foundation Meishan photograph ↗
  52. Field evidence for coal combustion links the 252 Ma Siberian Traps with global carbon disruption ↗
  53. Roger Smith on the Permian mass extinction event ↗
  54. Brink & Reisz 2014 ↗
  55. Pusch et al. 2024 ↗
  56. Maho et al. 2024 ↗
  57. Endothiodon histology and CT study ↗
  58. Cox & Angielczyk's feeding-system study ↗
  59. Rawson et al. 2024 ↗
  60. data.bris deposit ↗
  61. Huttenlocker & Botha-Brink 2014, histology ↗
  62. Duhamel et al. 2026 ↗
  63. Bajdek et al. 2016, coprolites ↗
  64. Benton 2021 synthesis ↗
  65. Knaus et al. 2021, femoral blood supply ↗
  66. Newham et al. 2020, cementum and blood flow ↗
  67. Meiri & Levin 2022 ↗
  68. authors' reply ↗
  69. Bolton, Mangera & Benoit 2025 ↗
  70. Benoit et al. 2026 ↗
  71. https://quentinmartinez.fr/wp-content/uploads/2023/07/Martinez_et_al_2023b.pdf ↗
  72. Day & Smith 2020, Endothiodon ↗
  73. Smith 2020, Cistecephalus ↗
  74. Viglietti 2020, Daptocephalus ↗
  75. Gastaldo and colleagues' 2020 climate-proxy study ↗
  76. Viglietti et al. 2017 ↗
  77. Smith and colleagues' 2021 study ↗
  78. Kammerer & Masyutin 2018 ↗
  79. Bulanov's 2024 Kotlassia distribution study ↗
  80. article deposit ↗
  81. Burgess & Bowring 2015 ↗
  82. 2025 review, The Anatomy and Lethality of the Siberian Traps Large Igneous Province ↗
  83. Grasby et al. 2011 ↗
  84. Augland et al. 2019 ↗
  85. DOI: 10.1038/s41467-026-74636-2 ↗
  86. Davydov 2021 ↗
  87. Callegaro et al. 2021 author manuscript ↗
  88. Zenodo model deposit ↗
  89. Sun et al. 2012, Lethally hot temperatures ↗
  90. Gliwa et al. 2022 ↗
  91. Brennecka et al. 2011 ↗
  92. Figshare 7357193 ↗
  93. Marquez et al. 2026 ↗
  94. GFZ's primary repository record ↗
  95. Sun et al. 2024 ↗
  96. https://historical-geobiology.stanford.edu/sites/g/files/sbiybj25131/files/media/file/penn_2018_science_hypoxia_and_end-permian_extinction.pdf ↗
  97. Author/museum PDF ↗
  98. Paper and supplement ↗
  99. Paper, CC BY ↗
  100. supplement ↗
  101. research team's account ↗
  102. 2022 polar wood study ↗
  103. Published paper ↗
  104. author PDF ↗
  105. author bibliography ↗
  106. author-deposited abstract and supplement listing ↗
  107. Viglietti et al. 2022 Dryad deposit ↗
  108. Original paper ↗
  109. https://doi.org/10.1130/0016-7606(1996)108%3C0195:GCEPME%3E2.3.CO;2 ↗
  110. Galesaurus study ↗
  111. Kulik et al. 2021 ↗
  112. Primary histology study ↗
  113. Smith & Botha 2005 ↗
  114. author preprint deposit ↗
  115. PeerJ paper ↗
  116. DOI: 10.1038/srep24053 ↗
  117. Study ↗
  118. Sidor et al. 2013 basin comparison ↗
  119. S2 NMQR 3636 STL ↗
  120. S3 BP/1/9332 ↗
  121. S4 BP/1/4011 ↗
  122. Jasinoski & Abdala 2017 ↗
  123. directory and License.txt ↗
  124. Study ↗
  125. 2023 tectonic/climate study ↗
  126. 2017 terrestrial Permian review, repository PDF ↗
  127. 2023 diversification analysis ↗
  128. Smithsonian specimen context ↗
  129. 2025 fusuline study ↗
  130. Groves & Altiner 2005 ↗
  131. Clapham & Bottjer 2007 assemblages ↗
  132. 2024 Jiantianba study ↗
  133. Publisher record ↗
  134. 2024 regional sponge-reef synthesis ↗
  135. NPS Guadalupe Mountains reef ↗
  136. Carlsbad geology ↗
  137. Paper, especially Fig. 12 ↗
  138. Fig. 10 SVG/PNG ↗
  139. Fig. 1 PNG ↗
  140. S5 comparisons ↗
  141. Fig. 2 ↗
  142. Primary paper ↗
  143. Nanjing University Meishan context ↗
  144. Smithsonian collection object ↗
  145. Permian-Triassic Mayhem ↗
  146. U53A-08, Trouble Upstairs ↗
  147. The Karoo Basin ↗
  148. Siberian Traps ↗
  149. Large igneous provinces lecture ↗
  150. Oldest Dad fossil discussion ↗

Music

  • Meditation From The Star Orion
    OctahedronMusic · Envato Elements
Footage and image usage

Credits and usage times retained from the published edition or its production credit record. Where present, times identify visual appearances, not documentary chapters. Source and licence corrections are listed in the visual credits above.

  • Anish Sheela — 6:49-6:59
  • Christian F. Kammerer — 5:20-5:25
  • Daderot — 4:36-4:40
  • Engin Akyurt — 11:44-11:55
  • Esv - Eduard Solà Vázquez — 17:06-17:12
  • European Space Agency — 9:45-9:49, 9:53-9:59
  • flowcomm — 1:43-1:53
  • Ghedoghedo — 4:14-4:26, 4:44-4:52, 14:49-14:52
  • James St. John — 8:30-8:34
  • Jeff Schmaltz, MODIS Land Rapid Response Team, NASA GSFC — 9:49-9:53
  • Jonathan Chen — 17:12-17:19
  • Nkansahrexford — 2:09-2:12, 2:17-2:21, 2:21-2:26, 14:59-15:04, 15:21-15:25
  • Scotese and Wright (2018) — 0:00-0:15, 1:22-1:31, 1:31-1:41, 9:12-9:21, 14:11-14:22, 14:40-14:46
  • The Trustees of the Natural History Museum, London — 4:52-4:57
  • Zachi Evenor — 4:06-4:14

How this film was made

Created and editorially directed by Anders Lunde using a custom production system developed over six months. AI assists research, drafting, programming, and visual production; I review and edit the material and make the final editorial decisions. Narration uses text-to-speech. Visuals combine custom-built scientific animations, AI-generated reconstructions, and credited scientific imagery.

Generated reconstructions illustrate scientific interpretations. They are distinct from photographs, specimens, and measured data.

More about the creator and production →