Episode 5.3 · 27 min

The Rise of Jawed Fishes

Changing Ordovician seas, mass extinction, and the fossil evidence for the rise of jawed fishes through the Silurian and Early Devonian.

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Visual credits

31 sources

Visual sources include third-party material incorporated into custom animations. Creator and licence details are included where documented; some source records remain incomplete. A missing licence is not a claim of public-domain status or permission to reuse the material.

  • Leptograptus fossil graptolites (Viola Formation, Ordovician; Murray County, Oklahoma, USA) (17391415055)

    James St. John

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Brachiopods in limestone (Richmondian Stage, Upper Ordovician; Southgate Hill roadcut, Indiana, USA) 1

    James St. John

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Amplexopora filiasa (fossil bryozoan) (Upper Ordovician; Cincinnati, Ohio, USA)

    James St. John

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Iocrinus subcrassus, crinoid, Late Ordovician, McMillan Formation, Butler County, Ohio, USA - Houston Museum of Natural Science - DSC01499

    Daderot

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Flexicalymene meeki (fossil trilobite) (Upper Ordovician; Cincinnati area, USA)

    James St. John

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Endoceras spp Svezia Museo Milano

    Antonov

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Sacabambaspis janvieri multiple specimens

    Ghedo

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Sacabambaspis janvieri cast

    Bloopityboop

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Arandaspis prionotolepis fossil

    Michal Maňas

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Arandaspis prionotolepis model P 231098

    Peter Roberts (model); Rodney Start (photograph); Museums Victoria

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Overview of Eriptychius americanus PF 1795

    Richard P. Dearden, Agnese Lanzetti, Sam Giles, Zerina Johanson, Andy S. Jones, Stephan Lautenschlager, Emma Randle & Ivan J. Sansom

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Tooth structures of the Panderodus specimen from the town of Waukesha

    Duncan J. E. Murdock, M. Paul Smith

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Conodont crystal orientation maps — Shirley et al. 2024 Figure 2

    Bryan Shirley and colleagues

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Body fossil of Panderodus from the waukesha biota

    Duncan J. E. Murdock & M. Paul Smith

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Ordovician glacial pavement, Tassili n'Ajjer

    IUGS International Commission on Geoheritage; individual photographer not established

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Scientific figure · source page 3

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

    Editorial use; no open license asserted.

  • 2026 fish genus-richness curve — standalone Figure 1A

    Wahei Hagiwara and Lauren Sallan

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Gotland-reef hg

    Hannes Grobe

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Qianodus holotype

    Plamsome

  • Tujiaaspis vividus — Nature Figure 1 holotype and anatomical drawing

    Zhikun Gai and colleagues; specimen photography Lijian Peng, drawing Aijuan Shi

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Cephalaspis Wiki1

    Petr Menshikov ( https://twitter.com/Petr75113553 ; https://vk.com/prehistoricproduction )

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Zhu et al. 2022 Figure 2 — Xiushanosteus mirabilis fossils and anatomical reconstruction

    Min Zhu and colleagues / IVPP

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

    Editorial use; no open license asserted.

  • Zhu et al. 2022 Figure 3 — Shenacanthus vermiformis fossil and interpretation

    You-an Zhu and colleagues / IVPP; Nature (Springer Nature)

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

    Editorial use; no open license asserted.

  • Eosteus chongqingensis — Nature Figure 1 fossil views

    You-an Zhu and colleagues

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

    No separate commercial reuse permission is recorded.

  • Megamastax amblyodus — Nature Figure 1 fossil and CT views

    Jing Lu and colleagues

    No separate commercial reuse permission is recorded.

  • Gemuendina stuertzi cast (cropped)

    Bloopityboop

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Doliodus pectoral skeleton and spine array — complete Figure 2

    Maisey and colleagues, American Museum of Natural History

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

    No separate commercial reuse permission is recorded.

  • Youngolepis praecursor Fig1

    Xindong Cui, Matt Friedman, Tuo Qiao, Yilun Yu & Min Zhu

  • Granddad, Shedd Aquarium, Chicago

    CusterDome

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Polypterus senegalus - Senegal-Flösselhecht, close up

    5snake5

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Dunkleosteus at Cleveland Museum of Natural History big

    Robosorne

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

Animation design references 5 sources

These works informed the design of explanatory animations. A reference listed here is not necessarily reproduced on screen.

  • Sacabambaspis tail specimens — Pradel et al. 2007 Figure 1

    Pradel and colleagues

  • Sacabambaspis revised tail interpretation — Pradel et al. 2007 Figure 2

    Pradel and colleagues

  • Scottish cephalaspids — Figure 5.4 A/B headshield drawings and C complete fossil

    Stensio 1932 / Natural History Museum London, reproduced in Scottish GCR

    Figure or page excerpt cropped from the source publication.

  • Gillis et al. 2013 Figure 5 — Dlx patterning and serial homology of pharyngeal arches

    Gillis, Modrell and Baker

    Figure or page excerpt cropped from the source publication.

  • Dunkleosteus terrelli 2024 reconstruction

    Russell K. Engelman

Research references

Selected sources from the episode’s research and scientific figures.

  1. ICS June 2026 chart ↗
  2. PaleoAtlas ↗
  3. Stigall et al., review and abstract ↗
  4. Servais and Harper, terminology review ↗
  5. paper PDF ↗
  6. Frey, USGS Professional Paper 1066-P ↗
  7. Song et al. 2026 ↗
  8. Klug et al. 2015, author-posted paper ↗
  9. 2025 size discussion ↗
  10. Mironenko, Endocerids: suspension feeding nautiloids? ↗
  11. Paper PDF ↗
  12. Pradel et al. 2007, tail and specimen figures ↗
  13. Ichnology, palaeoecology and taphonomy of a Gondwanan early vertebrate habitat ↗
  14. Animal history timescale, Arandaspis calibration ↗
  15. Exact collection record ↗
  16. Dearden et al. 2023 ↗
  17. Keating et al. 2018 ↗
  18. Sansom et al. 2005 ↗
  19. Brazeau et al. 2020 ↗
  20. Murdock and Smith 2021, version of record and figures ↗
  21. Balter et al. 2019, calcium isotopes ↗
  22. Primary paper ↗
  23. 2024 review ↗
  24. 2025 Earth-Science Reviews synthesis ↗
  25. Sheehan 2001 review ↗
  26. Harper 2024 review ↗
  27. Primary paper ↗
  28. Bond & Grasby 2020 ↗
  29. Primary paper ↗
  30. Global Hirnantian brachiopod review ↗
  31. Huang et al. 2018 recovery analysis ↗
  32. Gillis, Modrell and Baker 2013 ↗
  33. Skate pseudobranch paper ↗
  34. Gai et al./Tujiaaspis primary record ↗
  35. swimming study ↗
  36. Nature primary description ↗
  37. Nature primary description ↗
  38. Silurian Times 31, China research report ↗
  39. Zhu et al. 2022 primary description and figures ↗
  40. associated specimen CT data ↗
  41. Zhu et al. 2009 ↗
  42. Zhu et al. 2013 ↗
  43. 10.1126/science.aah3764 ↗
  44. 2023 post-thoracic study ↗
  45. 10.1126/sciadv.aeb2297 ↗
  46. Dryad associated data and code ↗
  47. Lu et al. 2016, Meemannia ↗
  48. Zhu, Wang & Fan 1994, Qujing assemblages ↗
  49. Henderson et al., 2023 review ↗
  50. Cui et al. 2022 ↗
  51. Sallan et al. 2018 ↗
  52. Campbellton ↗
  53. Qujing 2024 ↗
  54. Brazeau & Friedman 2015 ↗
  55. Tatsumi et al. 2016 ↗
  56. Zhang et al. 2023 ↗
  57. Li et al. 2025 ↗
  58. Bubphamanee et al. 2025, PNAS e2501342122 ↗
  59. 2023 size analysis ↗
  60. Engelman 2024, Palaeontologia Electronica ↗
  61. Anderson & Westneat 2007 ↗
  62. Lebedev et al. 2023 ↗
  63. 2025 review, Fig. 7 ↗
  64. 2026 Evolution study ↗
  65. Nature 2022 Fig. 2 ↗
  66. corporate purchase terms ↗
  67. Cincinnati Under the Sea: Ordovician Cephalopods ↗
  68. KpGAj8INgd8 ↗
  69. ICS December 2024 chart ↗
  70. model documentation ↗
  71. Rasmussen et al. 2016 ↗
  72. Algeo et al. 2016 review ↗
  73. Servais et al. 2016, institutional article record ↗
  74. Camp Nelson fossils, NPS, Emily Jackson photo credit ↗
  75. NPS fossil series ↗
  76. NPS fossil brachiopods ↗
  77. USGS Ordovician faunal monograph ↗
  78. 2025 study, searchable full-text record ↗
  79. institutional bibliographic record ↗
  80. Sea Animals & Their Descendants ↗
  81. Evolution & Extinction ↗
  82. museum diorama tour ↗
  83. Gagnier and Blieck 1992 ↗
  84. Canadian early-vertebrate synthesis ↗
  85. Young 2009, ANU record ↗
  86. Museums Victoria educational resource ↗
  87. Houée et al. 2023 histology ↗
  88. institutional PDF ↗
  89. Miyashita et al. 2019 ↗
  90. Goudemand et al. 2011, apparatus reconstruction ↗
  91. Soom Shale review, author-posted paper ↗
  92. Fig. 2 data ↗
  93. Houée et al. 2025 review ↗
  94. Jandzik et al. 2015 ↗
  95. Manchester manuscript ↗
  96. 2024 carbon-cycle modeling paper and stratigraphic summary ↗
  97. Rasmussen et al. 2023 reassessment ↗
  98. Kozik et al. 2022 AGU Advances ↗
  99. Primary article/repository ↗
  100. Figshare dataset ↗
  101. age framework ↗
  102. 2025 brachiopod analysis ↗
  103. JNCC British Silurian palaeontology ↗
  104. 2024 Gotland study ↗
  105. data service ↗
  106. https://paleobiodb.org/data1.2/): ↗
  107. Gillis et al. 2009 anatomical/developmental study ↗
  108. Primary abstract ↗
  109. Kuratani and colleagues' developmental account ↗
  110. 2021 cyclostome evo-devo review ↗
  111. zebrafish study ↗
  112. 2021 skate transcription study ↗
  113. 2020 skate study ↗
  114. 2022 swimming-speed comparative study ↗
  115. ncomms2429 PDF ↗
  116. Birmingham author PDF, including geological supplement ↗
  117. Birmingham author PDF ↗
  118. IVPP bibliographic record ↗
  119. Rongxi shallow-marine red-bed study ↗
  120. The 2019 scale study ↗
  121. Zhu et al. 2016, Science 354:334–336 ↗
  122. Bianchengichthys, 2021 ↗
  123. alternate repository ↗
  124. Birmingham institutional release ↗
  125. CAS image-search source ↗
  126. wius98NCkhM ↗
  127. 2026 Devonian actinopterygian jaw study ↗
  128. Johanson et al. 2006, Eoactinistia, primary paper text ↗
  129. Maisey et al. 2017, Doliodus, AMNH publication ↗
  130. Scottish Geological Conservation Review chapter 5 ↗
  131. Mesacanthus revision ↗
  132. Vaškaninová & Kraft 2014, *The largest Lower Devonian placoderm—Antineosteus rufus* ↗
  133. authors' Charles University record ↗
  134. Mark-Kurik & Carls 2004 ↗
  135. The 2024 marine megafauna review ↗
  136. Ferrón & Botella's thelodont ecological study ↗
  137. National Museums Scotland collection review ↗
  138. Kennedy et al. 2012 ↗
  139. Wang et al. 2024 ↗
  140. institutional abstract ↗
  141. figure record ↗
  142. AMNH Gemuendina cast photograph ↗
  143. Senckenberg Hunsrück collection ↗
  144. Ohio Geological Survey's 2025 GEOfacts sheet ↗
  145. 2025 exhibit explanation ↗
  146. Engelman 2023, Diversity 15:318 ↗
  147. *Giant, swimming mouths* ↗
  148. Gayford et al. 2024, *Cautionary tales on the use of proxies to estimate body size and form of extinct animals* ↗
  149. Anderson & Westneat 2007 ↗
  150. Smithsonian D. marsaisi collection record ↗
  151. 2017 Moroccan vertebrate review ↗
  152. Anatolepis reanalysis ↗

Music

  • Meditation From The Star Orion
    OctahedronMusic · Envato Elements

How this film was made

Created and editorially directed by Anders Lunde using a custom production system developed over six months. AI assists research, drafting, programming, and visual production; I review and edit the material and make the final editorial decisions. Narration uses text-to-speech. Visuals combine custom-built scientific animations, AI-generated reconstructions, and credited scientific imagery.

Generated reconstructions illustrate scientific interpretations. They are distinct from photographs, specimens, and measured data.

More about the creator and production →