Episode 5.3 · 27 min
The Rise of Jawed Fishes
Changing Ordovician seas, mass extinction, and the fossil evidence for the rise of jawed fishes through the Silurian and Early Devonian.
Watch this film on YouTube ↗Visual credits
31 sourcesSources and credits for imagery used in this film, including material incorporated into explanatory animations.
Leptograptus fossil graptolites (Viola Formation, Ordovician; Murray County, Oklahoma, USA) (17391415055)
James St. John
View original source ↗Source licence: CC BY 2.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Brachiopods in limestone (Richmondian Stage, Upper Ordovician; Southgate Hill roadcut, Indiana, USA) 1
James St. John
View original source ↗Source licence: CC BY 2.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Amplexopora filiasa (fossil bryozoan) (Upper Ordovician; Cincinnati, Ohio, USA)
James St. John
View original source ↗Source licence: CC BY 2.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Iocrinus subcrassus, crinoid, Late Ordovician, McMillan Formation, Butler County, Ohio, USA - Houston Museum of Natural Science - DSC01499
Daderot
View original source ↗Source licence: CC0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Flexicalymene meeki (fossil trilobite) (Upper Ordovician; Cincinnati area, USA)
James St. John
View original source ↗Source licence: CC BY 2.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Endoceras spp Svezia Museo Milano
Antonov
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Sacabambaspis janvieri multiple specimens
Ghedo
View original source ↗Source licence: CC BY-SA 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Sacabambaspis janvieri cast
Bloopityboop
View original source ↗Source licence: CC BY-SA 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Arandaspis prionotolepis fossil
Michal Maňas
View original source ↗Source licence: Public domainIncorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Arandaspis prionotolepis model P 231098
Peter Roberts (model); Rodney Start (photograph); Museums Victoria
View original source ↗Source licence: CC BY 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Overview of Eriptychius americanus PF 1795
Richard P. Dearden, Agnese Lanzetti, Sam Giles, Zerina Johanson, Andy S. Jones, Stephan Lautenschlager, Emma Randle & Ivan J. Sansom
View original source ↗Source licence: CC BY 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Tooth structures of the Panderodus specimen from the town of Waukesha
Duncan J. E. Murdock, M. Paul Smith
View original source ↗Source licence: CC BY-SA 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Conodont crystal orientation maps — Shirley et al. 2024 Figure 2
Bryan Shirley and colleagues
View original source ↗Source licence: CC BY 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Body fossil of Panderodus from the waukesha biota
Duncan J. E. Murdock & M. Paul Smith
View original source ↗Source licence: CC BY 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Ordovician glacial pavement, Tassili n'Ajjer
IUGS International Commission on Geoheritage; individual photographer not established
Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Scientific figure · source page 3
Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
2026 fish genus-richness curve — standalone Figure 1A
Wahei Hagiwara and Lauren Sallan
View original source ↗Source licence: CC BY 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Gotland-reef hg
Hannes Grobe
View original source ↗Source licence: CC BY-SA 2.5Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Qianodus holotype
Plamsome
View original source ↗Source licence: CC BY-SA 4.0Tujiaaspis vividus — Nature Figure 1 holotype and anatomical drawing
Zhikun Gai and colleagues; specimen photography Lijian Peng, drawing Aijuan Shi
Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Cephalaspis Wiki1
Petr Menshikov ( https://twitter.com/Petr75113553 ; https://vk.com/prehistoricproduction )
View original source ↗Source licence: CC BY-SA 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Zhu et al. 2022 Figure 2 — Xiushanosteus mirabilis fossils and anatomical reconstruction
Min Zhu and colleagues / IVPP
Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Zhu et al. 2022 Figure 3 — Shenacanthus vermiformis fossil and interpretation
You-an Zhu and colleagues / IVPP; Nature (Springer Nature)
Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Eosteus chongqingensis — Nature Figure 1 fossil views
You-an Zhu and colleagues
Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Megamastax amblyodus — Nature Figure 1 fossil and CT views
Jing Lu and colleagues
Gemuendina stuertzi cast (cropped)
Bloopityboop
View original source ↗Source licence: CC BY-SA 4.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Doliodus pectoral skeleton and spine array — complete Figure 2
Maisey and colleagues, American Museum of Natural History
Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Youngolepis praecursor Fig1
Xindong Cui, Matt Friedman, Tuo Qiao, Yilun Yu & Min Zhu
View original source ↗Source licence: CC BY 4.0Granddad, Shedd Aquarium, Chicago
CusterDome
View original source ↗Source licence: CC0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Polypterus senegalus - Senegal-Flösselhecht, close up
5snake5
View original source ↗Source licence: CC0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Dunkleosteus at Cleveland Museum of Natural History big
Robosorne
View original source ↗Source licence: CC BY-SA 3.0Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.
Animation design references 5 sources
These works informed the design of explanatory animations. A reference listed here is not necessarily reproduced on screen.
Sacabambaspis tail specimens — Pradel et al. 2007 Figure 1
Pradel and colleagues
Sacabambaspis revised tail interpretation — Pradel et al. 2007 Figure 2
Pradel and colleagues
Scottish cephalaspids — Figure 5.4 A/B headshield drawings and C complete fossil
Stensio 1932 / Natural History Museum London, reproduced in Scottish GCR
Figure or page excerpt cropped from the source publication.
Gillis et al. 2013 Figure 5 — Dlx patterning and serial homology of pharyngeal arches
Gillis, Modrell and Baker
Figure or page excerpt cropped from the source publication.
Dunkleosteus terrelli 2024 reconstruction
Russell K. Engelman
View original source ↗Source licence: CC BY 4.0
Research references
Selected sources from the episode’s research and scientific figures.
- ICS June 2026 chart ↗
- PaleoAtlas ↗
- Stigall et al., review and abstract ↗
- Servais and Harper, terminology review ↗
- paper PDF ↗
- Frey, USGS Professional Paper 1066-P ↗
- Song et al. 2026 ↗
- Klug et al. 2015, author-posted paper ↗
- 2025 size discussion ↗
- Mironenko, Endocerids: suspension feeding nautiloids? ↗
- Paper PDF ↗
- Pradel et al. 2007, tail and specimen figures ↗
- Ichnology, palaeoecology and taphonomy of a Gondwanan early vertebrate habitat ↗
- Animal history timescale, Arandaspis calibration ↗
- Exact collection record ↗
- Dearden et al. 2023 ↗
- Keating et al. 2018 ↗
- Sansom et al. 2005 ↗
- Brazeau et al. 2020 ↗
- Murdock and Smith 2021, version of record and figures ↗
- Balter et al. 2019, calcium isotopes ↗
- Primary paper ↗
- 2024 review ↗
- 2025 Earth-Science Reviews synthesis ↗
- Sheehan 2001 review ↗
- Harper 2024 review ↗
- Primary paper ↗
- Bond & Grasby 2020 ↗
- Primary paper ↗
- Global Hirnantian brachiopod review ↗
- Huang et al. 2018 recovery analysis ↗
- Gillis, Modrell and Baker 2013 ↗
- Skate pseudobranch paper ↗
- Gai et al./Tujiaaspis primary record ↗
- swimming study ↗
- Nature primary description ↗
- Nature primary description ↗
- Silurian Times 31, China research report ↗
- Zhu et al. 2022 primary description and figures ↗
- associated specimen CT data ↗
- Zhu et al. 2009 ↗
- Zhu et al. 2013 ↗
- 10.1126/science.aah3764 ↗
- 2023 post-thoracic study ↗
- 10.1126/sciadv.aeb2297 ↗
- Dryad associated data and code ↗
- Lu et al. 2016, Meemannia ↗
- Zhu, Wang & Fan 1994, Qujing assemblages ↗
- Henderson et al., 2023 review ↗
- Cui et al. 2022 ↗
- Sallan et al. 2018 ↗
- Campbellton ↗
- Qujing 2024 ↗
- Brazeau & Friedman 2015 ↗
- Tatsumi et al. 2016 ↗
- Zhang et al. 2023 ↗
- Li et al. 2025 ↗
- Bubphamanee et al. 2025, PNAS e2501342122 ↗
- 2023 size analysis ↗
- Engelman 2024, Palaeontologia Electronica ↗
- Anderson & Westneat 2007 ↗
- Lebedev et al. 2023 ↗
- 2025 review, Fig. 7 ↗
- 2026 Evolution study ↗
- Nature 2022 Fig. 2 ↗
- corporate purchase terms ↗
- Cincinnati Under the Sea: Ordovician Cephalopods ↗
- KpGAj8INgd8 ↗
- ICS December 2024 chart ↗
- model documentation ↗
- Rasmussen et al. 2016 ↗
- Algeo et al. 2016 review ↗
- Servais et al. 2016, institutional article record ↗
- Camp Nelson fossils, NPS, Emily Jackson photo credit ↗
- NPS fossil series ↗
- NPS fossil brachiopods ↗
- USGS Ordovician faunal monograph ↗
- 2025 study, searchable full-text record ↗
- institutional bibliographic record ↗
- Sea Animals & Their Descendants ↗
- Evolution & Extinction ↗
- museum diorama tour ↗
- Gagnier and Blieck 1992 ↗
- Canadian early-vertebrate synthesis ↗
- Young 2009, ANU record ↗
- Museums Victoria educational resource ↗
- Houée et al. 2023 histology ↗
- institutional PDF ↗
- Miyashita et al. 2019 ↗
- Goudemand et al. 2011, apparatus reconstruction ↗
- Soom Shale review, author-posted paper ↗
- Fig. 2 data ↗
- Houée et al. 2025 review ↗
- Jandzik et al. 2015 ↗
- Manchester manuscript ↗
- 2024 carbon-cycle modeling paper and stratigraphic summary ↗
- Rasmussen et al. 2023 reassessment ↗
- Kozik et al. 2022 AGU Advances ↗
- Primary article/repository ↗
- Figshare dataset ↗
- age framework ↗
- 2025 brachiopod analysis ↗
- JNCC British Silurian palaeontology ↗
- 2024 Gotland study ↗
- data service ↗
- https://paleobiodb.org/data1.2/): ↗
- Gillis et al. 2009 anatomical/developmental study ↗
- Primary abstract ↗
- Kuratani and colleagues' developmental account ↗
- 2021 cyclostome evo-devo review ↗
- zebrafish study ↗
- 2021 skate transcription study ↗
- 2020 skate study ↗
- 2022 swimming-speed comparative study ↗
- ncomms2429 PDF ↗
- Birmingham author PDF, including geological supplement ↗
- Birmingham author PDF ↗
- IVPP bibliographic record ↗
- Rongxi shallow-marine red-bed study ↗
- The 2019 scale study ↗
- Zhu et al. 2016, Science 354:334–336 ↗
- Bianchengichthys, 2021 ↗
- alternate repository ↗
- Birmingham institutional release ↗
- CAS image-search source ↗
- wius98NCkhM ↗
- 2026 Devonian actinopterygian jaw study ↗
- Johanson et al. 2006, Eoactinistia, primary paper text ↗
- Maisey et al. 2017, Doliodus, AMNH publication ↗
- Scottish Geological Conservation Review chapter 5 ↗
- Mesacanthus revision ↗
- Vaškaninová & Kraft 2014, *The largest Lower Devonian placoderm—Antineosteus rufus* ↗
- authors' Charles University record ↗
- Mark-Kurik & Carls 2004 ↗
- The 2024 marine megafauna review ↗
- Ferrón & Botella's thelodont ecological study ↗
- National Museums Scotland collection review ↗
- Kennedy et al. 2012 ↗
- Wang et al. 2024 ↗
- institutional abstract ↗
- figure record ↗
- AMNH Gemuendina cast photograph ↗
- Senckenberg Hunsrück collection ↗
- Ohio Geological Survey's 2025 GEOfacts sheet ↗
- 2025 exhibit explanation ↗
- Engelman 2023, Diversity 15:318 ↗
- *Giant, swimming mouths* ↗
- Gayford et al. 2024, *Cautionary tales on the use of proxies to estimate body size and form of extinct animals* ↗
- Anderson & Westneat 2007 ↗
- Smithsonian D. marsaisi collection record ↗
- 2017 Moroccan vertebrate review ↗
- Anatolepis reanalysis ↗
Music
- Meditation From The Star Orion
OctahedronMusic · Envato Elements
How this film was made
Created and editorially directed by Anders Lunde using a custom production system developed over six months. AI assists research, drafting, programming, and visual production; I review and edit the material and make the final editorial decisions. Narration uses text-to-speech. Visuals combine custom-built scientific animations, AI-generated reconstructions, and credited scientific imagery.
Generated reconstructions illustrate scientific interpretations. They are distinct from photographs, specimens, and measured data.
More about the creator and production →